Background: HIV cure requires the reactivation and elimination of latent reservoirs which persist in T cells and macrophages despite antiretroviral therapy (ART). However, cure trials to date have been hampered by an inability of the endogenous immune system to eliminate reactivated HIV+ cells. Immunotherapy strategies to augment NK cell-mediated elimination of HIV+ cells via antibody dependent cellular cytotoxicity (ADCC) may be useful to enhance HIV elimination in vivo.
Methods: NK cells from people with HIV (PWH) on suppressive ART and seronegative controls were immunophenotyped and their cytotoxic responses to HIV-infected T cells and monocyte derived macrophages assessed via CD107a degranulation. The ability of anti-HIV antibodies with reduced Fc fucosylation to elicit ADCC responses against HIV+ targets was also assessed.
Results: NK cells from PWH showed reduced ADCC activity towards both HIV+ T cells and macrophages as compared to HIV- controls (p<0.001 and 0.01, respectively). Immunophenotyping of degranulating NK cells identified an enhanced ADCC ability of NKG2A+ and KIR3DL2+ NK cells to target HIV+ T cells and macrophages, respectively (p<0.05 for both). However, in PWH the proportion of NKG2A+ NK cells was significantly reduced (19% vs 49% in controls, p=0.03). In contrast, memory-like NK cells (e.g. CD57+, FcRγ-) were expanded in PWH but showed impaired ADCC responses against both cell types. Fc-modification of anti-HIV envelope antibodies including NIH45-46 and 35O22 by afucosylation significantly enhanced ADCC responses of NK cells against HIV+ T cell and macrophage targets.
Conclusion: NK cells from PWH have an impaired ability to target HIV+ cells via ADCC, likely related to immune exhaustion and a skewing of NK cell populations towards memory-like cells with reduced anti-HIV ADCC potential. Enhancing the ADCC-eliciting potential of anti-HIV antibodies through afucosylation modifications offers the potential to overcome these deficits and enhance ADCC-mediated targeting of HIV+ cells in HIV cure approaches.